Some concerns I have regarding Lemmy and Fediverse architecture

So if I understand this right, you pick a server and your server account can post on that server and any servers that that server federates with.

So what happens to your account if the server you joined goes down? Yes, you could always create a new account somewhere else, but you lose all your followed communities and post and comment histories as well.

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You post and comment histories would likely still be saved on any instances that federated with them previously, I would imagine.

This is correct, I can find old posts from an account I deleted ages ago from an instance that doesn't exist still floating around those we federated with.

Well, if it was ages ago, instances may have gotten wise to the fact that your account is gone and deleted the posts by now.